Species account taken from: "A Revision of the Genus Ashmeadiella (Hymen., Megachilidae) Author(s): Charles D. Michener Source: American Midland Naturalist,Vol. 22, No. 1 (Jul., 1939), pp. 1-84"
Titusella pronitens Cockerell, 1906, Bull. Am. Mus. Nat. Hist., 22:446, 9 Cockerell, 1908, Ann. Mag. Nat. Hist., (8)2:331 (record); Cockerell, 1910, Univ. Colo. Stud., 7:191; Cockerell, 1919, Ann. Mag. Nat. Hist.; (9)4:358 (record); Cockerell, 1928, Univ. Colo. Stud., 16:115 (record); Cockerell, 1933, Ann. Ent. Soc. Am., 26:44 (record).
This species has the aspect and to a large degree the structure of cubiceps, to which it is closely related, differing primarily in the presence of a latge lobe between the first and second mandibular teeth.
Female: Inner margins of eyes diverging toward clypeus; under side of flagellum dusky; clypeus impunctate except laterally, anterior margin with median emnargination; mandibles with distance from first tooth to inner apical angle greater than distance from inner apical angle to base; angle between line drawn from third to fourth mandibular tooth and inner margin of mand- ibles about 1350; first mandibular tooth broadened basally on inner side into a rounded lobe, forming a distinct fifth tooth; cheeks much wider than eyes seen from side; supraclypeal area largely impunctate; frons and vertex closely punctate, posterior half of vertex more coarsely and less closely punctate than anterior part; anterior ocellus slightly anterior to midpoint between antennal bases and posterior edge of vertex; distance between posterior ocelli less than distance to eye, much less than distance to posterior edge of vertex. Scutum, except anteriorly, punctured like posterior part of vertex; tegulae dark brown; wings slightly dusky; both margins of inner hind tibial spur with about seven low, oblique teeth, outer margin with about ten similar teeth; mesepisterna slightly more finely punctate than scutum. Abdomen finely punctate, punc- tures of dorsum of tergites one to three so fine and sparse as to be incon- spicuous. Scopa yellowish white. Length 6I?2 to 8 mm.
This species occurs in the higher mountains of Colorado. Type: Female; Florissant, Colorado, in the Cockerell collection at Boulder, Colorado. Although in 1936 I considered this species a synonym of cubiceps, it now appears that it is distinct on the basis of the mandibular characters mentioned above. It may be only a subspecies of cubiceps, however, since morphologically it seems no more distinct from it than clypeata, which intergrades with cubicep