Cannabis sativa L.
Marijuana; Indian Hemp; Marihuana; Cannabis indica Lam; Cannabis ruderalis Janisch
Life   Plantae   Dicotyledoneae   Cannabaceae   Cannabis

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Cannabis sativa
© Copyright Malcolm Storey 2011-2118 · 3
Cannabis sativa
Cannabis sativa
© Copyright Malcolm Storey 2011-2118 · 3
Cannabis sativa

Cannabis sativa
© Copyright Malcolm Storey 2011-2118 · 3
Cannabis sativa
Cannabis sativa
© Copyright Malcolm Storey 2011-2118 · 3
Cannabis sativa

Cannabis sativa
© Copyright Malcolm Storey 2011-2118 · 3
Cannabis sativa
Cannabis sativa
© Copyright Malcolm Storey 2011-2118 · 3
Cannabis sativa

Cannabis sativa
© Copyright Malcolm Storey 2011-2118 · 3
Cannabis sativa
Cannabis sativa, seed
© Copyright Seeds USID 2015 · 1
Cannabis sativa, seed

Cannabis sativa, seed
© Copyright Seeds USID 2015 · 1
Cannabis sativa, seed
Cannabis sativa, F leaf flower
© Kay Yatskievych, 2003 · 1
Cannabis sativa, F leaf flower

Associates · map
FamilyScientific name @ source (records)
Agaricaceae  Phoma cannabis @ BPI (1)

Phoma glomerata @ BPI (1)
Botryosphaeriaceae  Botryosphaeria marconii @ BPI (8)

Lasiodiplodia theobromae @ BPI (1)

Phyllosticta cannabis @ BPI (8)
Chaetomiaceae  Chaetomium elatum @ BPI (1)
Cicadellidae  Ceratagallia ( @ UKYL_TCN (1)

Graphocephala coccinea @ UKYL_TCN (1)

Paraphlepsius irroratus @ UKYL_TCN (1)
Dermateaceae  Calloria luteola @ 802622B (1); 802622A (1)

Cylindrosporium @ BPI (1)
Diaporthaceae  Diaporthe tulasnei @ BPI (1)

Phomopsis cannabina @ BPI (4)

Phomopsis ganjae @ BPI (3)

Phomopsis @ BPI (1)
Erysiphaceae  Sphaerotheca macularis @ BPI (4)
Glomerellaceae  Colletotrichum coccodes @ BPI (1)

Colletotrichum dematium @ BPI (2)

Colletotrichum @ 396605A (1); 396605B (1)
Helotiaceae  Hymenoscyphus herbarum @ BPI (1)
Hypocreaceae  Hypomyces cancri @ BPI (1)
Mycosphaerellaceae  Cercospora canavaliae @ BPI (1)

Cercospora cannabina @ BPI (4)

Cercospora cannabis @ BPI (1)

Didymella cannabis @ 802692B (1); 802692A (1)

Pseudocercospora cannabina @ BPI (1)

Pseudocercospora cannabis @ BPI (1)

Ramularia collo-cygni @ BPI (1)

Septoria cannabina @ BPI (7)

Septoria cannabis @ 377728A (1); 377728B (1); BPI (49)
Nectriaceae  Fusarium graminearum @ BPI (1)

Fusarium ossicola @ 451243A (1); 451243B (1)

Fusarium oxysporum @ BPI (1)

Fusarium solani @ BPI (1)

Fusarium @ BPI (2)

Gibberella cyanogena @ BPI (1)

Gibberella quinqueseptata @ 632074B (1); 632074A (1)

Gibberella zeae @ BPI (1)
Peronosporaceae  Pseudoperonospora cannabina @ BPI (4)
Phaeosphaeriaceae  Leptosphaeria acuta @ BPI (1)

Ophiobolus anguillides @ BPI (2)
Pleosporaceae  Curvularia cymbopogonis @ BPI (1)

Leptosphaerulina trifolii @ BPI (1)

Stemphylium botryosum @ BPI (1)
Sclerotiniaceae  Botrytis cinerea @ BPI (1)

Botrytis @ BPI (2)
Tricholomataceae  Helotium herbarum @ BPI (1)
Turritellidae  Vermicularia dematium @ BPI (1)
Valsaceae  Diplodina cannabicola @ BPI (2)
Xylariaceae  Sphaeria cannabis @ BPI (1)
_  Ceratocapsidea complicata @ AMNH_PBI (1)

Coniothecium @ BPI (1)

Dendrophoma marconii @ BPI (4)

Macrophomina phaseolina @ BPI (3)

Micropeltopsis cannabis @ BPI (1)

Plenodomus cannabis @ BPI (2)

Torula herbarum @ BPI (1)

Trichothecium roseum @ BPI (3)

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Cannabis sativa L.

Syn.: Cannabis indica Lam.
Hemp, Marijuana, Cannabis oil

Source: James A. Duke. 1983. Handbook of Energy Crops. unpublished.

  1. Uses
  2. Folk Medicine
  3. Chemistry
  4. Toxicity
  5. Description
  6. Germplasm
  7. Distribution
  8. Ecology
  9. Cultivation
  10. Harvesting
  11. Yields and Economics
  12. Energy
  13. Biotic Factors
  14. References


A multiple-use plant, furnishing fiber, oil, medicine, and narcotics. Fibers are best produced from male plants. In the temperate zone, oil is produced from females which have been left to stand after the fiber-producing males have been harvested. Leaves are added to soups in southeast Asia. Varnish is made from the pressed seeds. Three types of narcotics are produced: hashish (bhang), the dried leaves and flowers of male and female shoots; ganja, dried unfertilized inflorescences of special female plants; and charas, the crude resin, which is probably the strongest. Modern medicine uses cannabis in glaucoma and alleviating the pains of cancer and chemotherapy. More resin is produced in tropical than in temperate climates. Lewis lung adenocarcinonoma growth has been retarded by oral administration of delta-9-tetrahydrocannabinol, delta-8-tetrahydrocannabinol and cannabinol, but not by cannabidiol. (J.N.C.I. 55: 597-602. 1975). The delta-9 also inhibits the replication of Herpes simplex virus.

Folk Medicine

Medicinally, plants are tonic, intoxicant, stomachic, antispasmodic, analgesic, narcotic, sedative and anodyne. Seeds and leaves are used to treat old cancer and scirrhous tumors. The seed, either as a paste or as an unguent, is said to be a folk remedy for tumors and cancerous ulcers. The decoction of the root is said help remedy hard tumors and knots in the joints. The leaf, prepared in various manners, is said to alleviate cancerous sores, scirrhous tumors, cold tumors, and white tumors. The plant is also used for mammary tumors and corns (C.S.I.R., 1948-1976). Europeans are said to use the dregs from Cannabis pipes in "cancer cures" (Watt and Breyer-Brandwijk, 1962). Few plants have a greater array of folk medicine uses: alcohol withdrawal, anthrax, asthma, blood poisoning, bronchitis, burns, catarrh, childbirth, convulsions, coughs, cystitis, delirium, depression, diarrhea, dysentery, dysmenorrhea, epilepsy, fever, gonorrhea, gout, inflammation, insomnia, jaundice, lockjaw, malaria, mania, mennorhagia, migraine, morphine withdrawal, neuralgia, palsy, rheumatism, scalds, snakebite, swellings, tetany, toothache, uteral prolapse, and whooping cough. Seeds ground and mixed with porridge given to weaning children.


Most varieties contain cannabinol and cannabinin; Egyptian variety contains cannabidine, cannabol and cannabinol, their biological activity being due to the alcohols and phenolic compounds. Resin contains crystalline compound cannin. Alcoholic extracts of American variety vary considerably in physiological activity. Per 100 g, the seed is reported to contain 8.8 g H 2 O, 21.5 g protein, 30.4 g fat, 34.7 g total carbohydrate, 18.8 g fiber, and 4.6 g ash. In Asia, per 100 g, the seed is reported to contain 421 calories, 13.6 g H 2 O, 27.1 g protein, 25.6 g fat, 27.6 g total carbohydrate, 20.3 g fiber, 6.1 g ash, 120 mg Ca, 970 mg P, 12.0 mg Fe, 5 mg beta-carotene equivalent, 0.32 mg thiamine, 0.17 mg riboflavin, and 2.1 mg niacin. A crystalline globulin has been isolated from defatted meal. It contains 3.8% glycocol, 3.6 alanine, 20.9 valine and leucine, 2.4 phenylalanine, 2.1 tyrosine, 0.3 serine, 0.2 cystine, 4.1 proline, 2.0 oxyproline, 4.5 aspartic acid, 18.7 glutamic acid, 14.4 tryptophane and arginine, 1.7 lysine, and 2.4% histidine. Oil from the seeds contains 15% oleic, 70% linoleic, and 15% linolenic and isolinolenic acids. The seed cake contains 10.8% water, 10.2% fat, 30.8% protein, 40.6% N-free extract, and 7.7% ash (20.3% K2O; 0.8% Na2O; 23.6% CaO, 5.7% MgO, 1.0% Fe2O3, 36.5% P2O5, 0.2% SO3; 11.9% SiO2, 0.1% Cl and a trace of Mn2O3). Trigonelline occurs in the seed. Cannabis also contains choline, eugenol, guaiacol, nicotine, and piperidine (C.S.I.R., 1948-1976), all listed as toxins by the National Institute of Occupational Safety and Health. A beta-resercyclic acid derivative has antibiotic and sedative properties; with a murine LD56 of 500 mg/kg, it has some aritiviral effect and inhibits the growth of mouse mammary tumor in egg embryo (Watt and Breyer-Brandwijk, 1962).


Non-users may suffer muscular incoordination (9 of 22 persons), dizziness (8), difficulty concentrating (8), confusion (7), difficulty walking (7), dysarthria (7), dry mouth (7), dysphagia (5), blurred vision (5), and vomiting (1), following oral ingestion of THC disguised in cookies (MMWR, October 20, 1978). People working with the plant or the fiber may develop dermatitis. In larger doses, hemp drugs may induce catalepsy, followed by coma and DEATH from cardiac failure (C.S.I.R., 1948-1976).


Annual herb, usually erect; stems variable, up to 5 m tall, with resinous pubescence, angular, sometimes hollow, especially above the first pairs of true leaves; basal leaves opposite, the upper leaves alternate, stipulate, long petiolate, palmate, with 3-11, rarely single, lanceolate, serrate, acuminate leaflets up to 10 cm long, 1.5 cm broad; flowers monoecious or dioecious, the male in axillary and terminal panicles, apetalous, with 5 yellowish petals and 5 poricidal stamens; the female flowers germinate in the axils and terminally, with one 1-ovulate ovary; fruit a brown, shining achene, variously marked or plain, tightly embracing the seed with its fleshy endosperm and curved embryo. Fl. summer; fr. late summer to early fall; year round in tropics. Seeds weigh 1.5-2.5 gm/100 seeds.


As Cannabis sativa has been cultivated for over 4,500 years for different purposes, many varieties and cultivars have been selected for specific purposes, as fiber, oil or narcotics. Drug-producing selections grow better and produce more drug in the tropics; oil and fiber producing plants thrive better in the temperate and subtropical areas. Many of the cultivars and varieties have been named as to the locality where it is grown mainly. However, all so called varieties freely interbreed and produce various combinations of the characters. The form of the plant and the yield of fiber from it vary according to climate and particular variety. Varieties cultivated particularly for their fibers have long stalks, branch very little, and yield only small quantities of seed. Varieties which are grown for the oil from their seed are short in height, mature early and produce large quantities of seed. Varieties grown for the drugs are short, much-branched with smaller dark-green leaves. Between these three main types of plants are numerous varieties which differ from the main one in height, extent of branching and other characteristics. Reported from the Central Asia, Hindustani, and Eurosiberian Centers of Diversity, marijuana or cvs thereof is reported to tolerate disease, drought, fungus, high pH, insects, laterite, low pH, mycobacteria, poor soil, slope, and weeds. (2 n = 20, 10, 40.)


Native to Central Asia, and long cultivated in Asia, Europe, and China. Now a widespread tropical, temperate and subarctic cultivar and waif. The oldest use of hemp seems to be for fiber, and later the seeds began to be used for culinary purposes. Plants yielding the drug seem to have been discovered in India, cultivated for medicinal purposes as early as 900 BC. In medieval times it was brought to North Africa where today it is cultivated exclusively for hashish or kif.


Plants very adaptable to soil and climatic conditions. Hemp for fiber requires a mild temperate climate with at least 67 cm annual rainfall, with abundant rain while seeds are germinating and until young plants become established. Thrives on rich, fertile, neutral to slightly alkaline, well-drained silt or clay loams with moisture retentive subsoils; does not grow well on acid, sandy soils. Of the many types of hemp, some are adapted to most vegetated terrains and climates. Ranging from Cool Temperate Steppe to Wet through Tropical Very Dry to Wet Forest Life Zones, marijuana is reported to tolerate annual precipitation of 3 to 40 dm (mean of 44 cases = 9.9 dm), annual temperature of 6 to 27°C (mean of 44 cases = 14.4), and pH of 4.5 to 8.2 (mean of 38 cases 6.5) (Duke, 1978, 1979).


Propagation mainly by seed. Experimentally, drug plants have been propagated from cuttings but such plants do not come true as to drug content of parent. Seeds stored in cool, dry place remain viable for up to two years. Hemp seed sown as early in spring as possible. Before sowing, land is plowed (in fall) several times to a depth of about 20-23 cm and repeatedly harrowed the land. In spring the land is harrowed again and rolled, making a firm tilth over the entire surface. In some areas a first plowing is done in the fall and red clover or lupin planted; in January or February a second plowing turns these under as a green-manure. Generally sown in March, seeds germinate at low temperature, but not below 1deg.C. Rate of seed sown varies with type of fiber desired; for coarse fiber for cordage and coarser textiles, 2.5 bu/ha is used; for finest fibers, 7.5-10 bu/ha used. Seed sown by machine in rows from 12 cm upwards, placing the seed at depth of 3.5 cm at rate of 40-60 kg/ha. In many countries seed sown broadcast. When grown for seed (oil), seed sown by drills; then such plants sometimes reach height of 5.3 m with thick stems up to 5 cm in diameter, much-branched. For fiber, stems up to 2 m tall and 0.5 cm in diameter are best; larger stems tend to get woody and have lower fiber content. Besides, they are more difficult to handle during harvesting, retting and scutching. Plants require little cultivation, except for weeding during early stages of growth. Hemp grows rapidly and soon crowds out weeds. After plants are 20 cm tall, weeding is abandoned. Hemp tends to exhaust the soil of nutrients. Some nutrients are returned to the soil after plants are harvested. On medium fertile soils a dressing of farm manure or a green-manure crop should be added and turned under. Chalk, potash, or gypsum may be applied to the soil to add the needed nutrition. Sodium nitrate and ammonium along with potassium sulfate have a beneficial effect on the fiber crop. Fiber-producing plants should always have plenty of proper nutrients, especially nitrogen, which is the most important element needed. Irrigation is seldom practiced.


Hemp is ready for harvest four to five months after planting, rarely earlier for some varieties. Harvesting depends on the climatic conditions, the variety of hemp grown and whether the crop is being grown for hemp or seed. In temperate areas, hemp is usually harvested from mid July to mid August. Both male and female plants look alike until they flower; then the male plants turn yellow and die, whereas the female plants remain dark green for another month until the seed ripens. Male plants are ready to harvest for fiber when the leaves change from dark green to light brown. The best yield of fiber (and only male plants are used) is then obtained. Hemp is harvested when the staminate flowers are beginning to open and shed their pollen. Seed is harvested from the female plants when most of it falls off when the plant is shaken. Best time of day to harvest seed is in early morning when fruits are turgid and conditions damp. As fruits dry out by mid-day, seed loss increases due to shattering. Usually stems are cut and the seeds shaken out over canvas sheets or beaten with sticks to extract the seeds. For fiber, hemp plants are cut by hand with a hemp knife, similar to a long-handled sickle. Plants are cut 2-3 cm above the ground and spread on the ground to dry. In some areas, the entire plants are pulled up and laid out to dry. Hand cutting, one man can cut about one-fifth hectare per day. Sometimes specially designed harvesters with a tractor are able to harvest four hectares a day. In many areas several varieties of hemp are grown so as to spread out the harvest, one maturing in late July and used later for seed crop in September, a second crop maturing in mid August, and a third maturing near end of August. Fiber is extracted from the stems of hemp by retting by methods similar to those used for other fiber plants. Sometimes the stems are dried before they are retted. After plants have air-dried for 4-6 days, the root and flower ends are cut off and the remaining portions, with branches and leaves taken or beaten off, are made into small bundles. For retting, 15-20 of these smaller bundles are made into larger bundles. In other areas stalks are not dried before retting, green stems, after roots and flower ends have been cut off, are made into bundles, and retted immediately. Hemp can be water retted, dew retted, or snow retted, according to the climatic conditions. The retted hemp stalks consist of fiber in the outer rind and a woody interior portion. Fiber is separated from the stalk by a breaking process. Stalks are dried after retting and the woody shive is broken into short pieces called hurds. Eventually the fibers are separated from the interior woody pieces by scutching by passing the bundles through a number of fluted rollers and then past large revolving drums with projecting bars which remove any remaining pieces of wood. Machines are able to handle 3-3.5 MT dried straw every hour, producing 0.4-0.5 MT of cleaned fiber.

Yields and Economics

Yields of hemp per hectare depend on climatic conditions, variety grown, soil and nutrition, and spacing of plants in the field. Weight of dried stems per hectare is usually between 4.5 to 7.5 T, with a yield of fiber about 25% of the dried stalks. Usually the taller the plant, the longer will be the fiber with a greater yield per plant. In some areas fiber yields of 850- 1,700 kg/ha compared to 1,300-1,700 kg/ha seed and 30 kg ganja. The U.S.S.R. is the largest producer of hemp in the world, producing about 33% of hemp fiber, annually 105,000 MT compared to the world production of 255,000 MT (excepting China). France and West Germany are the chief importers, Italy and Yugoslavia exporters. Chile, China, Japan and Peru also produce hemp. Narcotic production is usually clandestine, but there is legal marijuana production in India. India is the main producer and exporter of oil from the seed.


In India, plants remaining in the field after harvesting for fiber are allowed to set seed. They are cut after the fruits are ripened and dried and threshed for seed collection. Grown solely for seeds, an average crop yields 1.3 to 1.6 MT/ha seed. The world low production yield was 288 kg/ha in Democratic People's Republic of Korea, the international production yield was 613 kg/ha, and the world high production yield was 3,842 kg/ha in People's Republic of China.

Biotic Factors

Among diseases of Cannibis are: Botryosphaeria marconii (stem canker, wilt), Botrytis cinerea (gray mold), Cylindrosporium sp. (leaf spot), Fusarium sp. (canker, stem rot), Gibberella saubinetii (stem rot), Hypomyces cancri (?root rot), Macrophominia phaseoli, Phomopsis cannabina, Phymatotrichum omivorum (root rot), Sclerotinia sclerotiorum (stem rot, wilt), Sclerotium rolfsii (southern blight), Septoria cannabis (leaf spot). Nematodes include: Ditylenchus dipsaci, Heterodera humuli, Longidorus maximus, Meloidogyne hapla, M. incognita, M. incognita acrita, M. spp ., and Pratylenchus coffeae (Golden, p.c., 1984). Occasionally Orobanche ramosa is paratitic on the roots.


  • Duke, J.A. 1978. The quest for tolerant germplasm. p. 1-61. In: ASA Special Symposium 32, Crop tolerance to suboptimal land conditions. Am. Soc. Agron. Madison, WI.
  • Duke, J.A. 1979. Ecosystematic data on economic plants. Quart. J. Crude Drug Res. 17(3-4):91-110.
  • C.S.I.R. (Council of Scientific and Industrial Research). 1948-1976. The wealth of India. 11 vols. New Delhi.
  • Watt, J.M. and Breyer-Brandwijk, M.G. 1962. The medicinal and poisonous plants of southern and eastern Africa. 2nd ed. E.&S. Livingstone, Ltd., Edinburgh and London.
Complete list of references for Duke, Handbook of Energy Crops
Last update July 3, 1996

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Cannabis sativa L.

Image of Cannabis sativa

General Information
Symbol: CASA3
Group: Dicot
Family: Cannabaceae
Duration: Annual
Growth Habit : Forb/herb
Native Status : CAN   I
HI   I
L48   I
PR   I
Data Source and Documentation
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Plants-NRCS Logos
green round image for nativity Native blue round image for introduced Introduced ocre round image for introduced and nativity Both white round image for no status Absent/Unreported
image for native, but no county data Native, No County Data image for introduced, but no county data Introduced, No County Data both introduced and native, but no county data Both, No County Data
Native Status:
lower 48 status L48    Alaska status AK    Hawaii status HI    Puerto Rico status PR    Virgin Islands status VI    Navassa Island NAV    Canada status CAN    Greenland status GL    Saint Pierre and Michelon status SPM    North America NA   


click on a thumbnail to view an image, or see all the Cannabis thumbnails at the Plants Gallery

©Richard A. Howard. Provided by Smithsonian Institution, Richard A. Howard Photograph Collection . United States, MA, Bussey, Jamaica Plain. Usage Requirements .

©Robin R. Buckallew. United States, KS, Hutchinson. Usage Requirements .

Britton, N.L., and A. Brown. 1913. An illustrated flora of the northern United States, Canada and the British Possessions. 3 vols. Charles Scribner's Sons, New York. Vol. 1: 634. Provided by Kentucky Native Plant Society . Scanned by Omnitek Inc . Usage Requirements .




Click on a scientific name below to expand it in the PLANTS Classification Report.
Rank Scientific Name and Common Name
Kingdom Plantae – Plants
Subkingdom Tracheobionta – Vascular plants
Superdivision Spermatophyta – Seed plants
Division Magnoliophyta – Flowering plants
Class Magnoliopsida – Dicotyledons
Subclass Hamamelididae
Order Urticales
Family Cannabaceae – Hemp family
Genus Cannabis L. – hemp
Species Cannabis sativa L. – marijuana

Subordinate Taxa

The Plants Database includes the following 4 subspecies of Cannabis sativa . Click below on a thumbnail map or name for subspecies profiles. Plant is native (blue) Native Plant is introduced Introduced Plant is introduced Native and Introduced Related taxa legend Distribution of <i>
Cannabis sativa</i>
L. ssp. <i>
(Lam.) E. Small & Cronquist
Cannabis sativa ssp. indica
marijuana Distribution of <i>
Cannabis sativa</i>
L. ssp. <i>
sativa </i>
Cannabis sativa ssp. sativa
marijuana Distribution of <i>
Cannabis sativa</i>
L. ssp. <i>
var. <i>
sativa </i>
Cannabis sativa ssp. sativa var. sativa
marijuana Distribution of <i>
Cannabis sativa</i>
L. ssp. <i>
var. <i>
Cannabis sativa ssp. sativa var. spontanea

Legal Status

Noxious Weed Information
This plant is listed by the U.S. federal government or a state. Common names are from state and federal lists. Click on a place name to get a complete noxious weed list for that location, or click here for a composite list of all Federal and State Noxious Weeds .
Illinois marijuana Noxious weed
Minnesota hemp Prohibited noxious weed
Missouri marijuana Noxious weed
Pennsylvania marijuana Noxious weed
West Virginia marijuana Noxious weed
U.S. Weed Information
Cannabis sativa Mary Jane grass hashish hemp marijuana pot This plant can be weedy or invasive according to the authoritative sources noted below.This plant may be known by one or more common names in different places, and some are listed above. Click on an acronym to view each weed list, or click here for a composite list of Weeds of the U.S.
STATE Assorted authors. State noxious weed lists for 46 states . State agriculture or natural resource departments.
NE&GP Stubbendieck, J., G.Y. Friisoe,and M.R. Bolick. 1994. Weeds of Nebraska and the Great Plains . Nebraska Department of Agriculture, Bureau of Plant Industry. Lincoln, Nebraska.
SWSS Southern Weed Science Society. 1998. Weeds of the United States and Canada. CD-ROM . Southern Weed Science Society. Champaign, Illinois.
WSWS Whitson, T.D. (ed.) et al.. 1996. Weeds of the West . Western Society of Weed Science in cooperation with Cooperative Extension Services, University of Wyoming. Laramie, Wyoming.

Wetland Status

Interpreting Wetland Status

Related Links

More Accounts and Images
ARS Germplasm Resources Information Network (CASA3)
CalPhotos (CASA3)
Flora of North America (CASA3)
Integrated Taxonomic Information System (CASA3)
Jepson Interchange (University of California - Berkeley) (CASA3)
Native American Ethnobotany (University of Michigan - Dearborn) (CASA3)
USF Atlas of Florida Vascular Plants (CASA3)
University of Tennessee Herbarium (Distribution) (CASA3)
University of Wisconsin - Stevens Point Freckmann Herbarium (CASA3)
Related Websites
Atlas of Florida Vascular Plants: image (CASA3)
Industrial Hemp and Other Alternative Crops for Small Scale Tobacco Producers (CASA3)
NY-Cornell University: Poisonous Plant Databases (images) (CASA3)
North American Industrial Hemp Council (CASA3)
OR-Feasibility of Industrial Hemp Production in the Pacific Northwest (CASA3)
Poisonous Plants of North Carolina: abstract & image (CASA3)
USDA Farmers Bulletin No. 1935 (CASA3)
University of New Orleans: hemp line drawings (CASA3)



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FNA | Family List | FNA Vol. 3 | Cannabaceae | Cannabis

1. Cannabis sativa Linnaeus, Sp. Pl. 2: 1027. 1753.

Hemp, marihuana (marijuana), pot, grass, maryjane; chanvre, cannabis

Staminate plants usually taller, less robust than pistillate plants. Stems 0.2-6 m. Leaves: petioles 2-7 cm. Leaflet blades mostly 3-9, linear to linear-lanceolate, 3-15 × 0.2-1.7 cm, margins coarsely serrate; surfaces abaxially whitish green with scattered, yellowish brown, resinous dots, strigose, adaxially darker green with large, stiff, bulbous-based conic hairs. Inflorescences numerous. Flowers unisexual, often transitional flowers and flowers of opposite sex developing later. Staminate flowers: pedicels 0.5-3 mm; sepals ovate to lanceolate, 2.5-4 mm, puberulent; stamens caducous after anthesis, somewhat shorter than sepals; filaments 0.5-1 mm. Pistillate flowers ± sessile, enclosed by glandular, beaked bracteole and subtended by bract; perianth appressed to and surrounding base of ovary. Achenes white or greenish, mottled with purple, ovoid, somewhat compressed, 2-5 mm, with ± persistent perianth that sometimes flakes off. 2 n = 20.

Flowering early summer-fall; staminate plants generally dying after anthesis, pistillate plants remaining dark green, persisting until frost. Well-manured, moist farmyards, and in open habitats, waste places (roadsides, railways, vacant lots), occasionally in fallow fields and open woods; 0-2000 m; introduced; principal naturalized range (see map) Ont., Que.; Ark., Conn., Del., Ill., Ind., Iowa, Kans., Ky., Maine, Md., Mass., Mich., Minn., Mo., Nebr., N.H., N.J., N.Y., N.Dak., Ohio, Okla., Pa., R.I., S.Dak., Vt., Va., W.Va., Wis.; native to Asia.

Cannabis sativa has been reported as cultivated illegally and as apparently ruderal in all provinces and states except Alaska. It has been collected least frequently in Mississippi and Idaho. It seems to be best established in the prairies and plains of central North America.

Hemp is a short-day plant; flowering depends upon the latitude of origin. Races originating closer to the equator (and generally higher in psychointoxicant) require a longer induction period for flowering than races originating farther north.

The taxonomy of Cannabis sativa , a polymorphic species, has been debated in scientific and legal forums. The name C . sativa subsp. indica (Lamarck) E. Small & Cronquist has been applied to plants with a mean leaf content of the psychotomimetic (hallucinatory) delta-9-tetrahydrocannabinol of at least 0.3%; those with a lesser content fall under C . sativa subsp. sativa . When separate species are recognized, the name C . indica Lamarck has generally been applied to variants with high levels of the intoxicant chemical, whereas the name C . sativa Linnaeus, interpreted in a restricted sense, has generally been applied to plants selected for their yield of bast fibers in the stems. (The latter generally have taller, hollow stems with longer internodes and less branching than races selected for drug content.)

Superimposed on this dimension of variation is selection for nonabscising achenes in cultivation and abscising achenes in the wild (i.e., outside of cultivation). This is analagous to selection of nonshattering cereals from wild, shattering grasses. Achenes selected for cultivation tend to be longer than 3.8 mm and lack a basal constricted zone; by contrast, achenes selected for wild existence tend to be shorter than 3.8 mm and to have a basal constricted zone that seems to facilitate disarticulation and a mottled, persistent perianth apparently serving as camouflage.

Within Cannabis sativa subsp. sativa , the wild phase has been named C . sativa var. spontanea Vavilov (= C . ruderalis Janishevsky), in contrast to the domesticated C . sativa var. sativa . Within C . sativa subsp. indica , the wild phase (not to be expected in North America) has been designated C . sativa var. kafiristanica (Vavilov) E. Small & Cronquist, as distinct from the domesticated C . sativa var. indica . The chemical and morphologic distinctions by which Cannabis has been split into taxa are often not readily discernible, appear to be environmentally modifiable, and vary in a continuous fashion. For most purposes it will suffice to apply the name Cannabis sativa to all plants encountered in North America. *

The Iroquois used Cannabis sativa medicinally to convince patients that they had recovered. They also found it useful as a stimulant (D. E. Moerman 1986).

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Number of matches : 19
Query: SELECT * FROM img WHERE ready=1 and taxon like "Cannabis sativa%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

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Cannabis sativa
Cannabis sativa
Mary Jane
ID: 0000 0000 0105 1531 [detail]
© 2005 Louis-M. Landry

Cannabis sativa
Cannabis sativa
Mary Jane
ID: 0000 0000 0105 1532 [detail]
© 2005 Louis-M. Landry

Cannabis sativa
Cannabis sativa
Mary Jane
ID: 0000 0000 0105 1533 [detail]
© 2005 Louis-M. Landry

Cannabis sativa
Cannabis sativa
Mary Jane
ID: 0000 0000 0105 1534 [detail]
© 2005 Louis-M. Landry

Cannabis sativa
Cannabis sativa
ID: 0000 0000 0506 1328 [detail]
© 2006 Louis-M. Landry

Cannabis sativa
Cannabis sativa
ID: 0000 0000 0506 1329 [detail]
© 2006 Louis-M. Landry

Cannabis sativa
Cannabis sativa
ID: 0000 0000 0408 1472 [detail]
© 2008 Louis-M. Landry

Cannabis sativa
Cannabis sativa
ID: 0000 0000 0408 1473 [detail]
© 2008 Louis-M. Landry

Cannabis sativa
Cannabis sativa
ID: 0000 0000 0408 1474 [detail]
© 2008 Louis-M. Landry

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1117 1075 [detail]
© 2017 Thayne Tuason

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1117 1076 [detail]
© 2017 Thayne Tuason

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1217 1537 [detail]
© 2017 Barry Rice

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1217 1538 [detail]
© 2017 Barry Rice

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1217 1539 [detail]
© 2017 Barry Rice

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1217 1540 [detail]
© 2017 Barry Rice

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1217 1541 [detail]
© 2017 Barry Rice

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1217 1542 [detail]
© 2017 Barry Rice

Cannabis sativa
Cannabis sativa
ID: 0000 0000 1217 1543 [detail]
© 2017 Barry Rice

Cannabis sativa
Cannabis sativa
ID: 0000 0000 0518 0269 [detail]
© 2018 Dean Wm. Taylor

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