Sargent, Gard. & Forest. 8: 463. 1895.
Gray or Bebb's or long-beaked willow
Richardson in J. Franklin et al., Narr. Journey Polar Sea, 753. 1823, not Thuillier 1799;
(Rydberg) C. K. Schneider;
(Fernald) C. K. Schneider;
branches divaricate, sometimes ± brittle at base, yellow-brown to dark red-brown, not or weakly glaucous, pilose to glabrescent, peeled wood often with very dense striae, to 25 mm; branchlets yellow-green or red-brown, moderately to very densely villous to glabrescent.
stipules rudimentary or absent on early ones, apex acute, acuminate, or convex; petiole convex to flat adaxially, 2-5.5-13 mm, pubescent adaxially; largest medial blade narrowly oblong, narrowly elliptic, elliptic, oblanceolate, or obovate, 20-44-87 × 10-16-45 mm, base cuneate, convex, or rounded, margins flat, entire, crenate, or irregularly serrate, glands submarginal, apex acute, acuminate, or convex, abaxial surface glaucous, moderately densely pubescent or long-silky to glabrescent, hairs white or gray, wavy, adaxial finely impressed-reticulate, dull or slightly glossy, moderately densely pubescent, sparsely short-silky, or glabrescent, hairs white or gray; proximal blade margins entire, gland-dotted; juvenile blade yellowish green or reddish, pilose or sparsely to moderately densely tomentose or long-silky abaxially, hairs white.
staminate flowering just before leaves emerge, pistillate flowering as leaves emerge; staminate stout to globose, 10-42 × 7-16 mm, flowering branchlet 0.5-11 mm; pistillate loosely flowered, stout, slender, or subglobose, 16.5-85 × 9-32 mm, flowering branchlet 1-26 mm; floral bract tawny, 1.2-3.2 mm, apex rounded, abaxially hairy to glabrescent, hairs straight or wavy.
adaxial nectary oblong or ovate, 0.3-0.8 mm; filaments distinct or connate less than 1/2 their lengths, glabrous or hairy on proximal 1/2; anthers yellow or purple turning yellow, ellipsoid or shortly cylindrical, 0.5-0.8 mm.
adaxial nectary oblong or square, 0.3-0.8 mm; ovary obclavate, beak slightly bulged below styles (long-beaked); ovules 6-16 per ovary; styles 0.1-0.4 mm; stigmas slenderly to broadly cylindrical.
Flowering early Apr-late Jun. Riparian and upland conifer forests, wet lowland thickets,
treed bogs, stream margins, lakeshores, prairie margins, dry south-facing slopes, cienegas, seeps, disturbed areas; 0-3300 m; Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., P.E.I., Que., Sask., Yukon; Alaska, Ariz., Calif., Colo., Conn., Idaho, Ill., Ind., Iowa, Maine, Md., Mass., Mich., Minn., Mont., Nebr., Nev., N.H., N.J., N.Mex., N.Y., N.Dak., Ohio, Oreg., Pa., R.I., S.Dak., Utah, Vt., Wash., Wis., Wyo.; Asia.
occurs in Nunavut on Akimiski Island in James Bay.
forms natural hybrids with
S. candida, S. geyeriana, S. humilis,
. Reports of hybrids with
(C. K. Schneider 1921; M. L. Fernald 1950) are not based on convincing specimens, and synthetic hybrids could not be made (G. W. Argus 1974). Reports of hybrids with
(Fernald) are unverified. Controlled pollinations with
had low seed viability (A. Mosseler 1990).
Fernald, as species) is intermediate between parental species. It resembles
in having juvenile leaves densely woolly, mature leaves sparsely to moderately woolly abaxially, margins strongly revolute to crenulate, and ovaries woolly; and
in having stipes 2.8-3 mm and capsules long-beaked, 8-9 mm. The hybrid commonly occurs in Newfoundland.
: A plant with the pistillate catkins and flowers of
and the narrow, entire or slightly serrulate leaves with white and ferruginous hairs of
was collected by R. D. Dorn in a mixed population in Montana (Beaverhead County).
: Reported by C. K. Schneider (1921) and M. L. Fernald (1950) and successfully synthesized by G. W. Argus (1974, 1986).
is known from Ontario, based on an infertile pistillate specimen, and from Alberta and Saskatchewan, where it is relatively uncommon. It was successfully synthesized (G. W. Argus 1974, 1986) and controlled pollinations showed high seed viability (A. Mosseler 1990).